This article records new taxa of trace fossils of every kind that are scheduled to be described during the year 2019, as well as other significant discoveries and events related to trace fossil paleontology that are scheduled to occur in the year 2019.
Ivantsov, Nagovitsyn & Zakrevskaya (2019) describe traces of macroorganisms associated with the body imprints of trace-producers from Ediacaran deposits of the southeastern White Sea region (Russia).[1]
A study on the ichnotaxonMacaronichnus segregatis degiberti described on the basis of fossil burrows likely formed through selective sand feeding and excretion of polychaetes, aiming to reconstruct tracemaker behaviour, is published by Nara & Seike (2019).[3]
Horseshoe crab traces from the Middle Jurassic strata of the Imilchil area (Central High Atlas, Morocco), indicating the presence of horseshoe crabs at the southern margin of the Tethys Ocean, are described by Oukassou et al. (2019).[5]
Tracks of giant millipede-like arthropods, assigned to the ichnotaxon Diplichnites cuithensis and likely produced by members of the genus Arthropleura, are described from the Carboniferous (late Pennsylvanian) of the Graissessac Basin (southern France) by Moreau et al. (2019).[6]
Tracks produced by an edopoid temnospondyl are described from the Carboniferous (Viséan) Alston Formation (North Yorkshire, United Kingdom) by Bird et al. (2019), representing the stratigraphically oldest known tetrapod trackway from the United Kingdom reported so far and the oldest known record of Edopoidea.[7]
First tetrapod tracks from the Permian of Sardinia (Italy), assigned to the ichnogenusMerifontichnus and representing the oldest occurrence of the ichnogenus to date, are described by Citton et al. (2019).[8]
A study on the taphonomy of Permian (Cisuralian) tetrapod tracks from the Coconino and De Chelly formations of Arizona, and on the locomotion and phylogenetic relationships of the trackmakers, is published by Marchetti et al. (2019).[9]
Tetrapod tracks assigned to the ichnogenus Ichniotherium are reported from the eolian Coconino Sandstone by Francischini et al. (2019), who interpret these tracks as the oldest known evidence of occupation of deserts by non-amniote tetrapods (probably diadectomorphs.[10]
A revision of Lopingian tetrapod tracks from eolian paleoenvironments of Germany and Scotland is published by Marchetti, Voigt & Lucas (2019), who reject the interpretation of these tracks as monospecific associations of synapsid footprints, reporting them to be more diverse, and interpret them as evidence indicating that faunal turnover related to end-Guadalupianextinction event occurred not only at high-mid paleolatitudes, but also at low paleolatitudes of Pangaea.[11]
A revision of tetrapod tracks from the Upper Permian Val Gardena Formation of the Dolomites region in northern Italy is published by Marchetti, Voigt & Klein (2019), who name new ichnogenus Dolomitipes.[12]
A revision of tetrapod tracks from Permian-Early Triassic tracksites in the main Karoo Basin of South Africa is published by Marchetti et al. (2019), who name new ichnotaxonKaroopes gansfonteinensis.[13]
Tetrapod footprints, mostly quadruped trackways of large footprints most likely produced by kannemeyeriiformdicynodonts, are described from the Middle Triassic Cerro de las Cabras Formation (Argentina) by Lagnaoui et al. (2019), who name a new ichnotaxonPentasauropus argentinae.[14]
A new assemblage of lizard tracks, representing the largest such assemblage yet reported from the Cretaceous, is described from the Lower Cretaceous Jinju Formation (South Korea) by Kim et al. (2019), who name a new ichnotaxonNeosauroides innovatus.[16]
Complete step cycles produced by archosaurs, assigned to the ichnogenus Brachychirotherium, are described from the Upper Triassic Machraa Abbass Member of the Oued Oum Er Rbiaa Formation (Morocco) by Hminna et al. (2019), representing the first complete trackway of this ichnogenus in North Africa.[18]
A study on tracks assigned to the ichnogenus Chirotherium from the Middle Triassic Guanling Formation (Yunnan, China), produced by members of Archosauriformes, is published online by Xing & Klein (2019), who also describe the first tracks and trackways of the ichnogenus Rhynchosauroides from the Asian continent.[19]
Five invertebrate traces including a new ichnospecies of Diplichnites (Diplichnites rawi), were described from the latest Carboniferous of Shropshire, England by Hedge et al. (2019) [20]
Tetrapod burrow most likely produced by a notosuchiancrocodylomorph is described from the Upper Cretaceous Bauru Group (Brazil) by Martinelli et al. (2019).[21]
Costa-Pérez, Moratalla & Marugán-Lobón (2019) evaluate the utility of geometric morphometrics for determining the degree to which size, speed and taxonomy are contributing factors to the difference of bipedal dinosaur trackways.[23]
Small theropod footprints representing the ichnogenus Minisauripus, preserving high-definition skin traces, are described from the Lower Cretaceous Jinju Formation (South Korea) by Kim et al. (2019).[29]
A study on putative theropod footprints assigned to the ichnogenus Eubrontes is published by Weems (2019), who argues that bipedal sauropodomorphs were more likely trackmakers of these tracks.[30]
A study on sauropod tracks from the Jurassic Tafaytour tracksites (Argana Basin, Morocco), and on their implications for inferring forelimb posture in sauropod dinosaurs, is published by Lallensack et al. (2019).[31]
A study on sauropod tracks from a new dinosaur tracksite from the Middle JurassicIsli Formation (Morocco), providing evidence of presence of basaleusauropods in the Middle Jurassic-Early Cretaceous interval in the northwestern part of Gondwana, is published by Oukassou et al. (2019).[32]
A study on sauropod tracks from the Upper Jurassic Tianchihe Formation at the Guanxi site (Shanxi, China) is published online by Xing et al. (2019).[33]
An assemblage of sauropod tracks with a considerable size range, likely produced by an association of sauropods of different size-classes, is described from the Cretaceous Hengshan Formation (Zhejiang, China) by Xing et al. (2019).[34]
New sauropod trackway, representing the first record of a narrow-gauge sauropod trackway from the Cenomanian reported so far, will be described from the Candeleros Formation (Argentina) by Heredia et al. (2019).[35]
Probable track of a thyreophoran dinosaur, possibly produced by Isaberrysaura, is described from the Middle Jurassic Lajas Formation (Argentina) by Pablo et al. (2019).[37]
Ordovician arthropod trackways in the Borrowdale Volcanic Group were shown to be more likely to be death traces than evidence for early life on land by Shillito and Davies (2019). ,[38]
A study on non-avian dinosaur and bird tracks (representing some of the oldest known bird tracks) preserved in slabs used as building stones at the Chengde Mountain Resort, originating from the Tuchengzi Formation (China) and dating to the Jurassic-Cretaceous boundary, is published online by Xing et al. (2019).[39]
Description of non-avian dinosaur and bird tracks from the Upper Cretaceous Chignik Formation (southwestern Alaska), evaluating their implications for the knowledge of habitat preferences of northern high-latitude dinosaurs, is published by Fiorillo et al. (2019).[40]
Pterosaur tracks, larger than most other pterosaurian ichnites of the Late Jurassic age, are described from the Kimmeridgian carbonate deposits exposed in Wierzbica Quarry (Poland) by Elgh, Pieńkowski & Niedźwiedzki (2019).[43]
Evidence from coprolites, isolated worn teeth, fossil regurgitates and crushed or bite-marked dicynodont bones, indicating that Triassic archosaur Smok wawelski was at least an occasional osteophage consuming bones in a manner comparable to tyrannosaurid theropod dinosaurs, is presented by Qvarnström, Ahlberg & Niedźwiedzki (2019).[44]
A new late Pleistocene site containing bones and footprints of mammals is reported from Brazil by Oliveira et al. (2019).[45]
Large carnivore footprints, probably produced by Smilodon populator, will be described from a new ichnological site from the Late Pleistocene of Buenos Aires Province (Argentina) by Agnolin et al. (2019), who name a new ichnotaxonFelipeda miramarensis.[46]
A study on human footprints, handprints and other traces from the Upper Paleolithic of the Bàsura Cave (Italy), and on their implications for the knowledge of the behavior and social structure of the human group inhabiting this cave, is published by Romano et al. (2019).[48]
Trace fossils assigned to the ichnogenus Protopaleodictyon, of large size compared to other ichnospecies of this ichnogenus, are described from the CambrianStephen–Eldon formation transition (Alberta, Canada) by Morgan, Henderson & Pratt (2019), who name a new ichnospecies Protopaleodictyon aitkeni.[49]
A study on trace fossils from the Lower Triassic Dongchuan, Feixianguan and Jialingjiang formations (China), and on their implications for inferring how the Permian–Triassic extinction event affected the brackish-water ecosystem and how this ecosystem recovered in the Early Triassic, is published by Zhang et al. (2019).[50]
A study on trace fossils from the early Middle Triassic Luoping Biota (Yunnan Province, South China), and on their implications for inferring the timing of recovery of marine ecosystems after Permian–Triassic extinction event, is published by Luo et al. (2019).[51]
^Mostafa Oukassou; Abdelouahed Lagnaoui; André Charrière; Hafid Saber; Wahiba Bel Haouz; Gerard D. Gierliński; Hendrik Klein; Hassan Ibouh (2019). "New evidence of xiphosurids from the Middle Jurassic of Morocco: Palaeoenvironmental, palaeoecological and palaeobiogeographical implications". Palaeogeography, Palaeoclimatology, Palaeoecology. 516: 268–283. Bibcode:2019PPP...516..268O. doi:10.1016/j.palaeo.2018.11.034. S2CID133758009.
^Jean-David Moreau; Georges Gand; Emmanuel Fara; Jean Galtier; Nicolas Aubert; Stéphane Fouché (2019). "Trackways of Arthropleura from the Late Pennsylvanian of Graissessac (Hérault, southern France)". Historical Biology: An International Journal of Paleobiology. 33 (7): 996–1007. doi:10.1080/08912963.2019.1675055. S2CID208597626.
^Paolo Citton; Ausonio Ronchi; Simone Maganuco; Martina Caratelli; Umberto Nicosia; Eva Sacchi; Marco Romano (2019). "First tetrapod footprints from the Permian of Sardinia and their palaeontological and stratigraphical significance". Geological Journal. 54 (4): 2084–2098. Bibcode:2019GeolJ..54.2084C. doi:10.1002/gj.3285. S2CID135216684.
^Heitor Francischini; Spencer G. Lucas; Sebastian Voigt; Lorenzo Marchetti; Vincent L. Santucci; Cassandra L. Knight; John R. Wood; Paula Dentzien-Dias; Cesar L. Schultz (2019). "On the presence of Ichniotherium in the Coconino Sandstone (Cisuralian) of the Grand Canyon and remarks on the occupation of deserts by non-amniote tetrapods". PalZ. 94 (1): 207–225. doi:10.1007/s12542-019-00450-5. S2CID181356924.
^Lorenzo Marchetti; Sebastian Voigt; Spencer G. Lucas (2019). "An anatomy-consistent study of the Lopingian eolian tracks of Germany and Scotland reveals the first evidence of the end-Guadalupian mass extinction at low paleolatitudes of Pangea". Gondwana Research. 73: 32–53. Bibcode:2019GondR..73...32M. doi:10.1016/j.gr.2019.03.013. S2CID146780505.
^Lorenzo Marchetti; Sebastian Voigt; Hendrik Klein (2019). "Revision of Late Permian tetrapod tracks from the Dolomites (Trentino-Alto Adige, Italy)". Historical Biology: An International Journal of Paleobiology. 31 (6): 748–783. Bibcode:2019HBio...31..748M. doi:10.1080/08912963.2017.1391806. S2CID134771751.
^Abdelouahed Lagnaoui; Ricardo N. Melchor; Eduardo S. Bellosi; Pablo M. Villegas; Nahuel Espinoza; Aldo M. Umazano (2019). "Middle Triassic Pentasauropus-dominated ichnofauna from western Gondwana: Ichnotaxonomy, palaeoenvironment, biostratigraphy and palaeobiogeography". Palaeogeography, Palaeoclimatology, Palaeoecology. 524: 41–61. Bibcode:2019PPP...524...41L. doi:10.1016/j.palaeo.2019.03.020. S2CID133757156.
^Abdelkbir Hminna; Hendrik Klein; Tariq Zouheir; Abdelouahed Lagnaoui; Hafid Saber; Jens N. Lallensack; Mostafa Oukassou (2019). "The Late Triassic archosaur ichnogenus Brachychirotherium: first complete step cycles from Morocco, North Africa, with implications for trackmaker identification and ichnotaxonomy". Historical Biology: An International Journal of Paleobiology. 33 (5): 1–14. doi:10.1080/08912963.2019.1658097. S2CID202864611.
^Lida Xing; Hendrik Klein (2019). "Chirotherium and first Asian Rhynchosauroides tetrapod trackways from the Middle Triassic of Yunnan, China". Historical Biology: An International Journal of Paleobiology. 33 (6): 791–801. doi:10.1080/08912963.2019.1661409. S2CID202863096.
^Agustín G. Martinelli; Giorgio Basilici; Lucas E. Fiorelli; Carolina Klock; Joachim Karfunkel; Ariela Costa Diniz; Marcus V.T. Soares; André Marconato; João Ismael da Silva; Luiz Carlos B. Ribeiro; Thiago S. Marinho (2019). "Palaeoecological implications of an Upper Cretaceous tetrapod burrow (Bauru Basin; Peirópolis, Minas Gerais, Brazil)". Palaeogeography, Palaeoclimatology, Palaeoecology. 528: 147–159. Bibcode:2019PPP...528..147M. doi:10.1016/j.palaeo.2019.05.015. S2CID181458875.
^Yuong-Nam Lee; Hang-Jae Lee; Yoshitsugu Kobayashi; Ariana Paulina-Carabajal; Rinchen Barsbold; Anthony R. Fiorillo; Khishigjav Tsogtbaatar (2019). "Unusual locomotion behaviour preserved within a crocodyliform trackway from the Upper Cretaceous Bayanshiree Formation of Mongolia and its palaeobiological implications". Palaeogeography, Palaeoclimatology, Palaeoecology. 533: Article 109239. Bibcode:2019PPP...53309239L. doi:10.1016/j.palaeo.2019.109239. S2CID197584839.
^Matteo Belvedere; Diego Castanera; Christian A. Meyer; Daniel Marty; Octavio Mateus; Bruno Camilo Silva; Vanda F. Santos; Alberto Cobos (2019). "Late Jurassic globetrotters compared: A closer look at large and giant theropod tracks of North Africa and Europe". Journal of African Earth Sciences. 158: Article 103547. Bibcode:2019JAfES.15803547B. doi:10.1016/j.jafrearsci.2019.103547. S2CID198400391.
^Ricardo N. Melchor; David L. Rivarola; Aldo Martín Umazano; Magdalena Nalín Moyano; Fátima R. Mendoza Belmontes (2019). "Elusive Cretaceous Gondwanan theropods: the footprint evidence from central Argentina". Cretaceous Research. 97: 125–142. Bibcode:2019CrRes..97..125M. doi:10.1016/j.cretres.2019.01.004. S2CID133788331.
^Yuta Tsukiji; Yoichi Azuma; Fumito Shiraishi; Masateru Shibata (2019). "A diverse theropod footprint assemblage from the Lower Cretaceous Kitadani Formation, Tetori Group, central Japan". Cretaceous Research. 97: 16–33. Bibcode:2019CrRes..97...16T. doi:10.1016/j.cretres.2019.01.003. S2CID135285358.
^Jens N. Lallensack; Shinobu Ishigaki; Abdelouahed Lagnaoui; Michael Buchwitz; Oliver Wings (2019). "Forelimb orientation and locomotion of sauropod dinosaurs: insights from the ?Middle Jurassic Tafaytour tracksites (Argana Basin, Morocco)". Journal of Vertebrate Paleontology. 38 (5): e1512501. doi:10.1080/02724634.2018.1512501. S2CID92247018.
^Mostafa Oukassou; Hendrik Klein; Abdelouahed Lagnaoui; André Charrière; Hafid Saber; Gerard D. Gierliński; Jens N. Lallensack; Abdelkbir Hminna; Ayoub Boumaalif; Ahmed Oussou; Driss Ouarhache (2019). "Polyonyx-like tracks from Middle-?Upper Jurassic red beds of Morocco: implications for sauropod communities on southern margins of Tethys". Palaeogeography, Palaeoclimatology, Palaeoecology. 536: Article 109394. Bibcode:2019PPP...53609394O. doi:10.1016/j.palaeo.2019.109394. S2CID210315305.
^Lida Xing; Hendrik Klein; Enze Wang; W. Scott Persons IV (2019). "Upper Jurassic dinosaur tracks from the Tianchihe Formation of Shanxi Province, China: review and new observations". Historical Biology: An International Journal of Paleobiology. 33 (6): 1–8. doi:10.1080/08912963.2019.1661408. S2CID202857014.
^Lida Xing; Martin G. Lockley; Hendrik Klein; Chunyong Chou; Donghao Wang; W. Scott Persons IV (2019). "A mid-Cretaceous dinosaur track assemblage from the Hengshan Formation in central Zhejiang, China". Historical Biology: An International Journal of Paleobiology. 33 (7): 973–980. doi:10.1080/08912963.2019.1675053. S2CID208593638.
^Arturo Miguel Heredia; Pablo José Pazos; Diana Elizabeth Fernández; Ignacio Díaz Martínez; Marcos Comerio (2019). "A new narrow-gauge sauropod trackway from the Cenomanian Candeleros Formation, northern Patagonia, Argentina". Cretaceous Research. 96: 70–82. Bibcode:2019CrRes..96...70H. doi:10.1016/j.cretres.2018.11.016. hdl:11336/94924. S2CID134917409.
^Lida Xing; Martin G. Lockley; Hendrik Klein; Gerard D. Gierliński; Yong Ye; Jianping Zhang; W. Scott Persons IV; Tao Wang (2019). "First thyreophoran type tracks from the Middle Jurassic Chuanjie Formation of Yunnan Province, China". Ichnos: An International Journal for Plant and Animal Traces. 26 (1): 8–15. doi:10.1080/10420940.2018.1479962. S2CID134190731.
^J. Pazos Pablo; González Estebenet M. Candela; E. Cocca Sergio; Dania Pascua (2019). "The oldest record of a tyreophoran track in Gondwana: Geological implications of subaerial exposure in the lower part of the Lajas Formation at the Covunco section (Neuquén Basin), Patagonia, Argentina". Journal of South American Earth Sciences. 94: Article 102198. Bibcode:2019JSAES..9402198P. doi:10.1016/j.jsames.2019.05.014. S2CID181619510.
^Lida Xing; Martin G. Lockley; Tianming Du; Lijun Zhang; Hendrik Klein; Anthony Romilio; W. Scott Persons IV; Kuan Wang; Zhenyu Li; Xiaoqiao Wan (2019). "Dinosaur tracks from the Jurassic-Cretaceous boundary Tuchengzi Formation (Hebei Province, China) used as building stones in the Chengde imperial summer resort: age, ichnology, and history". Cretaceous Research. 107: Article 104310. Bibcode:2020CrRes.10704310X. doi:10.1016/j.cretres.2019.104310. S2CID210266977.
^Martin Ezequiel Farina; Verónica Krapovickas; Lucas Fernández Piana; Rocío Belen Vera; María De Los Ángeles Ordoñez (2019). "Flamingo-like footprints and the problem of addressing biological diversity in the past". Historical Biology: An International Journal of Paleobiology. 33 (7): 912–926. doi:10.1080/08912963.2019.1669024. S2CID208582248.
^Erik Elgh; Grzegorz Pieńkowski; Grzegorz Niedźwiedzki (2019). "Pterosaur track assemblages from the Upper Jurassic (lower Kimmeridgian) intertidal deposits of Poland: Linking ichnites to potential trackmakers". Palaeogeography, Palaeoclimatology, Palaeoecology. 530: 32–48. Bibcode:2019PPP...530...32E. doi:10.1016/j.palaeo.2019.05.016. S2CID182940454.
^Édison V. Oliveira; Pétrius S. Bélo; Gelson L. Fambrini; Alcides N. Sial; Ana K.B. Silva; Alcina M.F. Barreto (2019). "A new late Pleistocene ichnological site with mammal footprints from Brazil". Journal of South American Earth Sciences. 94: Article 102216. Bibcode:2019JSAES..9402216O. doi:10.1016/j.jsames.2019.102216. S2CID189964303.
^Federico L. Agnolin; Nicolás R. Chimento; Denise H. Campo; Mariano Magnussen; Daniel Boh; Francisco De Cianni (2019). "Large carnivore footprints from the Late Pleistocene of Argentina". Ichnos: An International Journal for Plant and Animal Traces. 26 (2): 119–126. Bibcode:2019Ichno..26..119A. doi:10.1080/10420940.2018.1479962. S2CID134190731.
^Chad A. Morgan; Charles M. Henderson; Brian R. Pratt (2019). "A giant Protopaleodictyon from the Middle Cambrian of western Canada". Ichnos: An International Journal for Plant and Animal Traces. 26 (3): 216–223. Bibcode:2019Ichno..26..216M. doi:10.1080/10420940.2018.1538981. S2CID135039835.
^Li-Jun Zhang; Luis A. Buatois; M. Gabriela Mángano; Yong-An Qi; Chao Tai (2019). "Early Triassic trace fossils from South China marginal-marine settings: Implications for biotic recovery following the end-Permian mass extinction". GSA Bulletin. 131 (1–2): 235–251. Bibcode:2019GSAB..131..235Z. doi:10.1130/B31867.1. S2CID135345831.