Simocyoninae is an extinct subfamily of Ailuridae (of which the only recent member is the red panda).[1] The taxonomic history of this group was complicated, as researchers placed various fossil caniform genera into the subfamily. In addition to Simocyon, there was also Oligobunis (an early mustelid), Cephalogale (a stem-bear), and Enhydrocyon (a hesperocyonine canid). This subfamily was initially classified within the Canidae or dog family. This idea went even further in 1910 when American paleontologist Henry Fairfield Osborn considered dholes, African wild dogs, and bush dogs to be the only extant representation left of the simocyonines. This was in large part to the overall similarity in the morphology of their molars, which suggested a shared ancestry of hypercarnivory. This view point was not supported by European paleontologists who believed that Simocyon was more closely related to musteloids. Soon the Simocyoninae were found as extinct subfamily of procyonids, or members of the raccoon-family, due to similarity of the basicranium of these animals. This was supported by subsequent researchers. It was not until a more completed skull of a Simocyon found in north Shaanxi, China was described by Wang (1997) who found that Simocyoninae is closely related Ailurinae on the basis of cranial and dental characteristics.[2]

Simocyoninae
Temporal range: Middle Miocene–Early Pliocene
Illustration of the skull from the genus Simocyon
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Ailuridae
Subfamily: Simocyoninae
Dawkins, 1868
Genera

Characteristics of the simocyonines that differentiate them from the ailurines are highly specialize premolars that enabled them to crush bone. This suggests the simocyonines were either specialized bone-crushing carnivorans like hyenas or were at least able to digest bone in their diet when consuming whole small animal prey. However it was not as fully developed as the aforementioned hyenas.[3]

References

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  1. ^ "†subfamily Simocyoninae". Palaeobiology Database.
  2. ^ Wang, X. (1997). "New cranial material of Simocyon from China, and its implications for phylogenetic relationship to the red panda (Ailurus)". Journal of Vertebrate Paleontology. 17 (1): 184–198. Bibcode:1997JVPal..17..184W. doi:10.1080/02724634.1997.10010963.
  3. ^ Peigné, S.; Salesa, M. J.; Antón, M.; Morales, J. (2005). "Ailurid carnivoran mammal Simocyon from the late Miocene of Spain and the systematics of the genus". Acta Palaeontologica Polonica. 50 (2): 219–238.