The El Mers Group is a geological group in the Middle Atlas of Morocco. It is subdivided into three formations named the El Mers I, II, and III Formations, respectively. It is a marine deposit primarily consisting of marl, with gypsum present in the upper part of unit 3. It is the lateral equivalent of the terrestrial Guettioua Sandstone. Dinosaur remains are among the fossils that have been recovered from the group, most notably those of sauropods and the unusual thyreophorans Adratiklit, Thyreosaurus, and Spicomellus.

El Mers Group
Stratigraphic range: BajocianCallovian
~168–166 Ma
TypeGroup
Sub-unitsEl Mers Formations 1–3
UnderliesUnconformity with Barremian or Plio-Pleistocene sediments
OverliesIch Timellaline–Bou Akrabene Formation
ThicknessOver 500 m (1,600 ft)
Lithology
PrimaryMarl, gypsum (only in unit 3)
OtherSandstone, limestone
Location
Coordinates33°30′N 4°18′W / 33.5°N 4.3°W / 33.5; -4.3
Approximate paleocoordinates28°18′N 2°00′W / 28.3°N 2.0°W / 28.3; -2.0
RegionBoulemane and Azilal Provinces
Country Morocco
ExtentMiddle Atlas
El Mers Group is located in Morocco
El Mers Group
El Mers Group (Morocco)

Description and paleoenvironment

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The El Mers formations are composed primarily of marls with intermittent layers of limestone and sandstone, characterized by abundant organic matter and benthic fauna.[1] It is delineated into five distinct members, each exhibiting diverse lithological characteristics. The three members comprise a combination of layers of marls, sandstones, and limestones, marked by various unique fossils and organic materials. Sedimentary discontinuities reveal deltaic sequences that exhibit varying sedimentary structures and lithological compositions, reflecting dynamic depositional environments ranging from subtidal to continental. The facies associations within the El Mers Formation are organized into elementary and mesosequences, delineating the transition from prodelta to deltaic plain environments. These sequences capture the evolution of tidal regimes and depositional dynamics.[2]

The shifts in facies and microfauna evolution in our study area reveal intriguing patterns, from the establishment of initial supratidal environments characterized by charophytic marls and ostracod limestones followed by a resurgence of marine conditions, evident from the reappearance of coastal facies and the diversification of ostracod faunas, indicating a significant transgressive phase during the Upper Bathonian–Lower Callovian period. While in the Central High Atlas marine influences ceased by the Upper Bajocian–Lower Bathonian giving way to continental red layers, in the Middle Atlas marine influences persisted until the Upper Bathonian–Lower Callovian boundary, indicating a diachronic filling of the Atlas gulfs towards the Tethyan domain from west to east.[3]

Fossils

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Color key
Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Notes
Uncertain or tentative taxa are in small text; crossed out taxa are discredited.

Dasycladales

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Genus Species Stratigraphic position Material Notes Images
Heteroporella[3]
  • H. lusitanica
  • SW part of the Skoura syncline
Calcareous imprints A dasycladacean algae of the family Dasycladaceae
Sarfiatella[3]
  • S. dubari
  • SW part of the Skoura syncline
Calcareous imprints A dasycladacean algae, associated in adjacent regions with Aalenian–Bajocian faunas

Foraminifera

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Genus Species Stratigraphic position Material Notes Images
Bullopora[3]
  • B. tuberculata
  • SW part of the Skoura syncline
Calcareous imprints A benthic foraminiferan of the family Polymorphinidae
Pfenderella[3]
  • SW part of the Skoura syncline
Calcareous imprints A benthic foraminiferan of the family Pfenderininae
Praekurnubia[3]
  • P. crusei
  • SW part of the Skoura syncline
Calcareous imprints A benthic foraminiferan of the family Pfenderinidae; a genus found in the Tethys between Bajocian and Oxfordian-aged layers
Pseudocyclammina[3]
  • P. maynci
  • SW part of the Skoura syncline
Calcareous imprints A benthic foraminiferan of the family Hauraniidae

Ostracoda

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Genus Species Stratigraphic position Material Notes Images
Cytherella[3]
  • C. index
  • Ait Hammou
Valves A freshwater/brackish ostracod of the family Cytherellidae
Fastigatocythere[3]
  • F. aff. brentonensis
  • Ait Hammou
Valves A freshwater/brackish ostracod of the family Progonocytheridae
Kinkelinella[3]
  • K. aff. triangula
  • Ait Hammou
Valves A freshwater ostracod of the family Progonocytheridae
Metacypris[3]
  • M. spp.
  • Ait Hammou
Valves A freshwater ostracod of the family Limnocytheridae. Local members of the genus genus would not be confined to limnic environments, but would be euryhaline or brackish, since they disappear when faunal diversity is at its maximum.
Procytheridea[3]
  • P. ihopyensis
  • Ait Hammou
Valves A freshwater ostracod of the family Schulerideidae

Xiphosura

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Genus Species Stratigraphic position Material Notes Images
Selenichnites[4]
  • S. tesiltus
  • S. isp.
  • Skoura Syncline
  • Taferdouste
Isolated crescent-shaped trace fossils Traces of xiphosurans, representing the second occurrence of this ichnogenus in Morocco

Mollusca

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Genus Species Stratigraphic position Material Notes Images
Cadomites[3]
  • C. cf. bremeri
  • East Skoura syncline
Isolated shells An ammonite of the family Stephanoceratidae and indicator of the Bathonian age
 
Example of specimens
Choffatia[5]
  • C. cf. urinacensis
  • East Skoura syncline
Isolated shells An ammonite of the family Perisphinctidae and indicator of the Callovian age
Clydoniceras[5][6]
  • C. discus
  • Tamguert n'Tarit
  • Tizi n'Juillerh
Isolated shells An ammonite of the family Clydoniceratidae
Modiolus[6]
  • M. imbricatus
  • El Mers
  • Tamguert n'Tarit
Isolated shells A marine mussel of the family Mytilidae
 
Example of extant member of the genus
Morphoceras[5]
  • M. macrescens
  • East Skoura syncline
Isolated shells An ammonite of the family Morphoceratidae and indicator of the Bathonian age
Oraniceras[5]
  • O. sp.
  • East Skoura syncline
Isolated shells An ammonite of the family Parkinsoniidae and indicator of the Bathonian age
Ostrea[5][6]
  • O. sandalina
  • El Mers
  • Tamguert n'Tarit
Isolated shells A marine oyster of the family Ostreidae
 
Example of extant member of the genus
Parkinsonia[3]
  • P. sp.
  • Skoura syncline
Isolated shells An ammonite of the family Parkinsoniidae and indicator of the Bajocian age for the lower portions of the Group
 
Example of specimen
Protocardia[5][6]
  • P. tikechkachensis
  • El Mers
  • Tamguert n'Tarit
Isolated shells A marine cockle of the family Cardiidae

Fishes

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Genus Species Stratigraphic position Material Notes Images
Lepidotes[7]
  • L. spp.
  • Ait Hammou
  • Djmila
  • Oued El Mers
  • Tizi n'Juillerh
Cranial material, isolated teeth and scales A marine/brackish bony fish of the family Lepidotidae
 
Example of specimen of the genus
Scheenstia[7]
  • S. cf. mantelli
  • Oued El Mers
Partial specimen wirth cranial material A marine/brackish bony fish of the family Lepidotidae
 
Reconstruction

Testudinata

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Genus Species Stratigraphic position Material Notes Images
Testudinata[5][6]
  • Indeterminate
  • Tamguert n'Tarit
  • Tizi n'Juillerh
Very numerous debris: escutcheons, plates, vertebrae, ribs, long bones Turtle remains of uncertain affinity

Crocodylomorpha

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Genus Species Stratigraphic position Material Notes Images
Hatcherichnus[8]
  • H. ispp.
  • Inzar O'Founass
Footprints Indeterminate Crocodrylomorph Tracks

"Megalosaurus"[5][6]

  • "M." mersensis
  • Botane
  • Tizi n'Juillerh

Isolated teeth and vertebra

A marine crocodylomorph of the family Teleosauridae. Mistaken as Theropod remains in the past.[9]

Steneosaurus?[5]

  • S.? spp.
  • Bou Iferaoun
  • Botane
  • Darak
  • Djmila
  • Oued Tamemecht
  • Tamguert r'Tane
  • Tirardine
  • Tissenfelt

Isolated teeth, cranial and postcranial material, both isolated and semi-articulated

A marine crocodylomorph of the family Teleosauridae. The referral to the genus Steneosaurus should be taken with caution, as this genus was overlumped.

Dinosauria

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Theropoda

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Genus Species Stratigraphic position Material Notes Images
Carmelopodus[8]
  • cf.C. isp.
  • Tassmante O'Moche
Footprints Theropod tracks of uncertain affinity, suggested to come from Ceratosauria members
Grallator?[8]
  • G.? isp.
  • Ifri N'Tfrane
Footprints Theropod Tracks, referred usually to small-medium sized Genera
Hispanosauropus?[8]
  • H.? isp.
  • Ifri N'Tfrane
Footprints Theropod tracks of uncertain affinity, different from Megalosauripus in the larger metatarsophalangeal region
Megalosauripus[10][8]
  • M. ispp.
  • cf.M. ispp.
  • El Mers
  • Tasra
  • Tasra Westbank
  • Tassmante O'Moche
  • Oued Tamghilt
Footprints Traces of theropods; members of the ichnofiber family Eubrontidae, incertae sedis within Theropoda
Averostra[5][10]
  • Indeterminate
  • Oued Botane
Isolated Teeth Isolated teeth of unidentified or undescribed/studied theropods. Teeth associated with the Teleosaur "Megalosaurus" mersensis have recently been suggesting to come from indeterminate Averostrans. Other remains need revision.[11][12]
Theropodipedia[8][13][10]
  • Indeterminate
  • El Mers
  • Inzar O'Founass
  • Ifri N'Tfrane
  • Laach O'Medda
  • Oulad Ali
  • Tasra
  • Tasra Westbank
  • Tassmante O'Moche
Footprints Theropod tracks of uncertain affinity, suggested to come from "allosaurs or megalosaurs"[13]

Sauropoda

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Genus Species Stratigraphic position Material Notes Images
Breviparopus[10][8]
  • B. isp.
  • El Mers
  • Oued Tamghilt
Footprints Traces of sauropods typical of the ichnofamily Parabrontopodidae, incertae sedis within Sauropodomorpha. Includes traces with pes similar to Diplodocoidea.[10]

Cetiosaurus[5][14]

  • "C." mogrebiensis
  • Aîn ou N'Jourh
  • Taghrout
  • Tamguert r'Tane
  • Tamguert n'Tarit
  • Tich Niouine
  • Oued Botane

Complete skeleton and referred co-types of partially complete and incomplete specimens

A eusauropod of the family Cetiosauridae, currently awaiting revision and likely not a member of the genus Cetiosaurus[15]

Sauropoda[16][17]
  • Indeterminate
  • Aït Tamjout
  • Boulahfa
  • 1 km east of Taguelft
Isolated and articulated bones Indeterminate or undescribed sauropod remains
Sauropodina[8][13][18]
  • Indeterminate
  • El Mers
  • Tasra
  • Tasra Westbank
  • Oulad Ali
Footprints Sauropod tracks of uncertain affinity. There are both small and enormous sauropod tracks, including some among the largest known worldwide, with a pes track length of up to 130 cm.[8]

Ornithischia

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Genus Species Stratigraphic position Material Notes Images
Adratiklit[19]
  • A. boulahfa
  • Boulahfa
NHMUK PV R37366 (A dorsal vertebra), three cervical vertebrae (NHMUK PV R37367; R37368), a dorsal vertebra (NHMUK PV R37365), and a left humerus (NHMUK PV R37007) A dacentrurine stegosaur  
Ornithopodichnus?[8]
  • O.? ispp.
  • Ifri N'Tfrane
Footprints Ornithichian Tracks, referred to short-toed forms such as Gilmoreosaurus
Spicomellus[20]
  • S. afer
  • Boulahfa
NHMUK PV R37412, a partial rib bearing four co-ossified spines An early ankylosaurian  
Thyreosaurus[21]
  • T. atlasicus
  • Boulahfa North
HIIUC-BN00, a partial postcranial skeleton that includes nine dorsal vertebrae and 21 dorsal rib remains, a limb bone (fibula?), and six dermal elements A dacentrurine stegosaur with unusual recumbent osteoderms  

Charophyta

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Genus Species Stratigraphic position Material Notes Images
Porochara[3]
  • P. hians
  • Ait Hammou
Oogonia A characean algae of the family Porocharaceae; charophytes are abundant in lower levels, associated with freshwater ostracods
 
Example of modern characean algae

Plantae

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Genus Species Stratigraphic position Material Notes Images
Protocupressinoxylon[22]
  • P. choubertii
  • Skoura syncline
Fossil wood Conifer wood of the family Hirmeriellaceae, considered to be a potential synonym of the genus Brachyoxylon
 
Example of wood trunk of the same genus

See also

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References

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  1. ^ Oukassou, M.; Boumir, Kh.; Benshili, Kh.; Ouarhache, D.; Lagnaoui, A.; Charrière, A. (June 2019). "The Tichoukt Massif: a Geotouristic Play in the Folded Middle Atlas (Morocco)". Geoheritage. 11 (2): 371–379. Bibcode:2019Geohe..11..371O. doi:10.1007/s12371-018-0287-y. ISSN 1867-2477. S2CID 134800520.
  2. ^ Soufiani, N.; Fedan, B. (2002). "Les dépôts bathoniens du synclinal d'El Mers (Moyen Atlas, Maroc): unités formationnelles, analyse et cadre géodynamique" (PDF). Bulletin de l'Institut scientifique. 24 (6): 1–14.
  3. ^ a b c d e f g h i j k l m n o Charrière, André; Dépêche, Françoise; Feist, Monique; Grambast-Fessard, Nicole; Jaffrezo, Michel; Peybernès, Bernard; Ramalho, Miguel (1994). "Microfaunes, microflores et paléoenvironnements successifs dans la formation d'El Mers (Bathonien-?Callovien) du synclinal de Skoura (Moyen-Atlas, Maroc)". Geobios. 27 (2): 157–174. Bibcode:1994Geobi..27..157C. doi:10.1016/s0016-6995(94)80002-2. ISSN 0016-6995.
  4. ^ Oukassou, Mostafa; Charrière, André; Lagnaoui, Abdelouahed; Gibb, Stacey; Michard, André; Saddiqi, Omar (April 2016). "First occurrence of the Ichnogenus Selenichnites from the Middle Jurassic Strata of the Skoura Syncline (Middle Atlas, Morocco); Palaeoecological and palaeoenvironmental context". Comptes Rendus Palevol. 15 (5): 461–471. Bibcode:2016CRPal..15..461O. doi:10.1016/j.crpv.2015.09.013.
  5. ^ a b c d e f g h i j k Lapparent, A. F. d. (1955). "Étude paléontologique des vertébrés du Jurassique d'El Mers (Moyen Atlas) [Paleontological study of the Jurassic veretebrates from El Mers (Middle Atlas)]". Notes et Mémoires du Service Géologique du Maroc. 124 (1): 1–36.
  6. ^ a b c d e f Termier, H.; Gubler, J.; Lapparent, A. F. (1940). "Reptiles et Poissons du Bathonien d'El-Mers (Moyen-Atlas marocain) [Reptiles and fishes from the Bathonian of El Mers (Moroccan Middle Atlas)]" (PDF). Comptes Rendus de l'Académie des Sciences de Paris. 210 (3): 768–770.
  7. ^ a b Beltan, Laurence (1957-01-01). "Etude d'un neurocrane de Lepidotes du Bathonien du Maroc". Bulletin de la Société Géologique de France. S6-VII (8–9): 1091–1106. doi:10.2113/gssgfbull.s6-vii.8-9.1091. ISSN 1777-5817.
  8. ^ a b c d e f g h i j Amzil, Mustapha; Oukassou, Mostafa; Lallensack, Jens N.; Klein, Hendrik; Zafaty, Omar; Saber, Hafid; Charrière, André; Meyer, Christian; Gierliński, Gerard D. (2024). "New dinosaur tracks from the Middle Jurassic red beds of the Middle Atlas (Morocco): Application of photogrammetry to ichnology and conservation of geological heritage". Proceedings of the Geologists' Association. 22 (3): 12–31. doi:10.1016/j.pgeola.2024.06.004.
  9. ^ Chabli, S. (1986). "Données nouvelles sur un «Dinosaurien» jurassique moyen du Maroc: Megalosaurus mersensis Lapparent 1955, et sur les Megalasaurides en Sénégal". Les dinosaures de la Chine à la France. 1 (2): 65–72.
  10. ^ a b c d e Hadri, M.; Lorente, F. P. (2012). "Historia de yacimientos con huellas de dinosaurio, desde su descubrimiento hasta su primer estudio: alrededores de El Mers (Marruecos)" (PDF). Zubia. 30 (6): 93–147.
  11. ^ "Non-theropods". theropoddatabase.github.io. Retrieved 2024-10-05.
  12. ^ "Neotheropoda". theropoddatabase.github.io. Retrieved 2024-11-01.
  13. ^ a b c Jenny, Jacques; Le Marrec, Alain; Monbaron, Michel (1981). "Les empreintes de pas de Dinosauriens dans le Jurassique moyen du Haut Atlas central (Maroc): Nouveaux gisements et precisions stratigraphiques". Geobios. 14 (3): 427–431. Bibcode:1981Geobi..14..427J. doi:10.1016/s0016-6995(81)80186-6. ISSN 0016-6995.
  14. ^ Allain, Ronan; Aquesbi, Najat (2008). "Anatomy and phylogenetic relationships of Tazoudasaurus naimi (Dinosauria, Sauropoda) from the late Early Jurassic of Morocco" (PDF). Geodiversitas. 30 (2): 345–424.
  15. ^ Läng, E. (2008). "Les Cétiosaures (Dinosaura, sauropoda) et les sauropodes du Jurassique moyen: revision systématique, nouvelles découvertes et implications phylogénétiques". (Doctoral dissertation, Paris, Muséum national d'histoire naturelle): 1–223. S2CID 160327994.
  16. ^ Termier, Henri (1942-01-01). "Donnees nouvelles sur le Jurassique rouge a dinosauriens du Grand et du Moyen-Atlas (Maroc)". Bulletin de la Société Géologique de France. S5-XII (4–6): 199–207. doi:10.2113/gssgfbull.s5-xii.4-6.199. ISSN 1777-5817.
  17. ^ Charroud, M.; Fedan, B. (1992). "Données préliminaires sur la découverte du gisement de Boulahfa à dinosauriens (SW de Boulemane, Moyen Atlas central)" (PDF). Notes et Mémoire du Service géologique du Maroc. 366 (5): 448–449.
  18. ^ Meyer, C. A.; Thüring, B. (2005). "Mind the Middle Jurassicgap-bone versus track record in dinosaurs". Abstracts 3rd Swiss Geoscience Meeting Zurich. 3 (1): 59–60.
  19. ^ Maidment, Susannah C.R.; Raven, Thomas J.; Ouarhache, Driss; Barrett, Paul M. (January 2020). "North Africa's first stegosaur: Implications for Gondwanan thyreophoran dinosaur diversity". Gondwana Research. 77: 82–97. Bibcode:2020GondR..77...82M. doi:10.1016/j.gr.2019.07.007. hdl:10141/622706.
  20. ^ Maidment, Susannah C. R.; Strachan, Sarah J.; Ouarhache, Driss; Scheyer, Torsten M.; Brown, Emily E.; Fernandez, Vincent; Johanson, Zerina; Raven, Thomas J.; Barrett, Paul M. (2021-09-23). "Bizarre dermal armour suggests the first African ankylosaur". Nature Ecology & Evolution. 5 (12): 1576–1581. Bibcode:2021NatEE...5.1576M. doi:10.1038/s41559-021-01553-6. ISSN 2397-334X. PMID 34556830. S2CID 237616095.
  21. ^ Zafaty, O.; Oukassou, M.; Riguetti, F.; Company, J.; Bendrioua, S.; Tabuce, R.; Charrière, A.; Pereda-Suberbiola, X. (2024). "A new stegosaurian dinosaur (Ornithischia: Thyreophora) with a remarkable dermal armour from the Middle Jurassic of North Africa". Gondwana Research. 131 (In press): 344–362. Bibcode:2024GondR.131..344Z. doi:10.1016/j.gr.2024.03.009. {{cite journal}}: |last4= has generic name (help)
  22. ^ Philippe, M.; Bamford, M.; McLoughlin, S.; Alves, L.S.R.; Falcon-Lang, H.J.; Gnaedinger, S.; Ottone, E.G.; Pole, M.; Rajanikanth, A.; Shoemaker, R.E.; Torres, T.; Zamuner, A. (2004). "Biogeographic analysis of Jurassic–Early Cretaceous wood assemblages from Gondwana". Review of Palaeobotany and Palynology. 129 (3): 141–173. Bibcode:2004RPaPa.129..141P. doi:10.1016/j.revpalbo.2004.01.005. ISSN 0034-6667.