Ursus ingressus (the Gamssulzen Cave bear) is an extinct species of the family Ursidae that lived in Central Europe during the Late Pleistocene. It is named after the Gamssulzen Cave in Austria, where the holotype of this species was found.[2]
| Ursus ingressus | |
|---|---|
Scientific classification 
| |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Carnivora |
| Family: | Ursidae |
| Genus: | Ursus |
| Species: | †U. ingressus
|
| Binomial name | |
| †Ursus ingressus Rabeder et al., 2004[1]
| |
Description
editUrsus ingressus was a large cave bear with massive, bulky limbs. It was larger than Ursus spelaeus, which has been estimated to weigh an average of 350 to 600 kg (770 to 1,320 lb) (male specimen).[2][3]
Behaviour
editSome studies have suggested the Gamssulzen Cave bear to have been herbivorous, living off vegetation with little contribution of grass.[4] Other studies proposed Ursus ingressus to have been an omnivore, with participation of terrestrial and more likely aquatic animal protein, that exceeds the participation of animal protein in the diet of the modern brown bear (Ursus arctos).[5] However it has also been suggested, that the feeding habits of cave bears can vary heavily depending on the environment.[6]
Skulls with bite damage from Zoolithen Cave in Germany suggest that Ursus ingressus came into conflicts with other big carnivores of the Late Pleistocene of Europe like the cave lion (Panthera spelaea) or the cave hyena (Crocuta crocuta spelaea).[7]
Distribution and habitat
editIt has been suggested that the Gamssulzen Cave bear dominated Ursus spelaeus in Central and Eastern Europe, while being out-competed by this species in Western Europe. Around 50,000 years ago, the Gamssulzen Cave bear migrated into the Alps and replaced two former populations of Ursus spelaeus: Ursus spelaeus eremus and Ursus spelaeus ladinicus. Ursus ingressus has been found as far east as the Ural Mountains in Russia and as far west as the Swabian Jura in Germany.[6][8]
It has mostly been found in medium and high elevated regions and probably was adapted to continental environments with cold and arid climate.[9]
Evolution and extinction
editUrsus ingressus and Ursus spelaeus evolved from Ursus deningeri and probably diverged between 173,000 and 414,000, or possibly as much as 600,000 years ago.[6][2] Some studies still question whether U. ingressus and U. spelaeus are separate species, instead treating them as subspecies of a single species.[9]
The Gamssulzen Cave bear survived U. spelaeus for about 1000 to 2000 years, locally replacing this species, but also became extinct about 30,000 years ago, just prior to the Last Glacial Maximum. The reasons for their extinction are still being discussed, with climate change and human hunting suggested as possible reasons.[8][10]
References
edit- ^ Rabeder, G., Hofreiter, M., Nagel, D., & Withalm, G. (2004). New taxa of alpine cave bears (Ursidae, Carnivora). Cahiers scientifiques-Muséum d'histoire naturelle de Lyon, 49-67.
- ^ a b c Gernot Rabeder (2004): Die Höhlenbären der Sulzfluh-Höhlen, Vorarlberger Naturschau 15, Seite 103–114
- ^ Per Christiansen (1999): What size were Arctodus simus and Ursus spelaeus (Carnivora: Ursidae)?, Finnish Zoological and Botanical Publishing Board, Helsinki 1999
- ^ Münzel, Susanne C. (2014). "Behavioural ecology of Late Pleistocene bears (Ursus spelaeus, Ursus ingressus): Insight from stable isotopes (C, N, O) and tooth microwear". Quaternary International. 339–340: 148–163. Bibcode:2014QuInt.339..148M. doi:10.1016/j.quaint.2013.10.020.
- ^ Dotsika, Elisavet (2011). "Palaeoclimatic information from isotopic signatures of Late Pleistocene Ursus ingressus bone and teeth apatite (Loutra Arideas Cave, Macedonia, Greece)". Quaternary International. 245 (2): 291–301. Bibcode:2011QuInt.245..291D. doi:10.1016/j.quaint.2011.01.027.
- ^ a b c Knapp, Michael (March 2009). "First DNA sequences from Asian cave bear fossils reveal deep divergences and complex phylogeographic patterns". Molecular Ecology. 18 (6): 1225–1238. doi:10.1111/j.1365-294x.2009.04088.x. PMID 19226321. S2CID 881987.
- ^ Diedrich, Cajus (2014). "Holotype skulls, stratigraphy, bone taphonomy and excavation history in the Zoolithen Cave and new theory about Esper's "great deluge"". E&G Quaternary Science Journal. 63: 78–98. doi:10.3285/eg.63.1.05. hdl:11858/00-1735-0000-0023-B183-5.
- ^ a b Münzel, Susanne C. (2011). "Pleistocene bears in the Swabian Jura (Germany): Genetic replacement, ecological displacement, extinctions and survival" (PDF). Quaternary International. 245 (2): 225–237. Bibcode:2011QuInt.245..225M. doi:10.1016/j.quaint.2011.03.060.
- ^ a b Baca, Mateusz, Stankovic, Anna, Stefaniak, Krzysztof, Marciszak, Adrian, Hofreiter, Michael, Nadachowski, Adam, Węgleński, Piotr, and Mackiewicz, Paweł. 2012. Genetic analysis of cave bear specimens from Niedźwiedzia Cave, Sudetes, Poland. Palaeontologia Electronica Vol. 15, Issue 2;21A,16p
- ^ Kitawaga, Keiko (September 2015). "Exploring cave use and exploitation among cave bears, carnivores and hominins in the Swabian Jura, Southwestern Germany". Journal of Taphonomy. 10 (Issue 3-4): 439–461.