The Cetaceans Portal
Cetacea (/sɪˈteɪʃə/; from Latin cetus 'whale', from Ancient Greek κῆτος (kêtos) 'huge fish, sea monster') is an infraorder of aquatic mammals belonging to the order Artiodactyla that includes whales, dolphins and porpoises. Key characteristics are their fully aquatic lifestyle, streamlined body shape, often large size and exclusively carnivorous diet. They propel themselves through the water with powerful up-and-down movement of their tail which ends in a paddle-like fluke, using their flipper-shaped forelimbs to maneuver.
While the majority of cetaceans live in marine environments, a small number reside solely in brackish water or fresh water. Having a cosmopolitan distribution, they can be found in some rivers and all of Earth's oceans, and many species inhabit vast ranges where they migrate with the changing of the seasons.
Cetaceans are famous for their high intelligence, complex social behaviour, and the enormous size of some of the group's members. For example, the blue whale reaches a maximum confirmed length of 29.9 meters (98 feet) and a weight of 173 tonnes (190 short tons), making it the largest animal ever known to have existed.
There are approximately 89 living species split into two parvorders: Odontoceti or toothed whales (containing porpoises, dolphins, other predatory whales like the beluga and the sperm whale, and the poorly understood beaked whales) and the filter feeding Mysticeti or baleen whales (which includes species like the blue whale, the humpback whale and the bowhead whale). Despite their highly modified bodies and carnivorous lifestyle, genetic and fossil evidence places cetaceans as nested within even-toed ungulates, most closely related to hippopotamus within the clade Whippomorpha. (Full article...)
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Kogia pusilla is an extinct species of sperm whale from the Middle Pliocene of Italy, also known as the Littleness Sperm Whale. related to the modern-day dwarf sperm whale (K. sima) and pygmy sperm whale (K. breviceps). It is known from a single skull discovered in 1877, and was considered a species of beaked whale until 1997. The skull shares many characteristics with other sperm whales, and is comparable in size to that of the dwarf sperm whale. Like the modern Kogia, it probably hunted squid in the twilight zone, and frequented continental slopes. The environment it inhabited was likely a calm, nearshore area with a combination sandy and hard-rock seafloor. K. pusilla likely died out due to the ice ages at the end of the Pliocene. (Full article...) -
Right whales are three species of large baleen whales of the genus Eubalaena: the North Atlantic right whale (E. glacialis), the North Pacific right whale (E. japonica) and the Southern right whale (E. australis). They are classified in the family Balaenidae with the bowhead whale. Right whales have rotund bodies with arching rostrums, V-shaped blowholes and dark gray or black skin. The most distinguishing feature of a right whale is the rough patches of skin on its head, which appear white due to parasitism by whale lice. Right whales are typically 13–17 m (43–56 ft) long and weigh up to 100 short tons (91 t; 89 long tons) or more.
All three species are migratory, moving seasonally to feed or give birth. The warm equatorial waters form a barrier that isolates the northern and southern species from one another although the southern species, at least, has been known to cross the equator. In the Northern Hemisphere, right whales tend to avoid open waters and stay close to peninsulas and bays and on continental shelves, as these areas offer greater shelter and an abundance of their preferred foods. In the Southern Hemisphere, right whales feed far offshore in summer, but a large portion of the population occur in near-shore waters in winter. Right whales feed mainly on copepods but also consume krill and pteropods. They may forage the surface, underwater or even the ocean bottom. During courtship, males gather into large groups to compete for a single female, suggesting that sperm competition is an important factor in mating behavior. Gestation tends to last a year, and calves are weaned at eight months old. (Full article...) -
Acrophyseter is a genus of extinct sperm whale that lived in the Late Miocene off the coast of what is now Peru. The genus comprises two species: A. deinodon and A. robustus. It is part of a group of macroraptorial sperm whales that all share several features for hunting large prey, such as deeply rooted and thick teeth. Acrophyseter measured 4–4.5 metres (13–15 ft) in length, making it the smallest macroraptorial sperm whale currently known. Because of its short pointed snout and strongly curved front teeth, it probably fed on the marine vertebrates of its time, such as seals and other whales. (Full article...) -
Livyatan is an extinct genus of macroraptorial sperm whale containing one known species: L. melvillei. The genus name was inspired by the biblical sea monster Leviathan, and the species name by Herman Melville, the author of the famous novel Moby-Dick about a white bull sperm whale. Herman Melville often referred to whales as "Leviathans" in his book. It is mainly known from the Pisco Formation of Peru during the Tortonian stage of the Miocene epoch, about 9.9–8.9 million years ago (mya); however, finds of isolated teeth from other locations such as Chile, Argentina, the United States (California), South Africa and Australia imply that either it or a close relative survived into the Pliocene, around 5 mya, and may have had a global presence. It was a member of a group of macroraptorial sperm whales (or "raptorial sperm whales") and was probably an apex predator, preying on whales, seals and so forth. Characteristically of raptorial sperm whales, Livyatan had functional, enamel-coated teeth on the upper and lower jaws, as well as several features suitable for hunting large prey.
Livyatan's total length has been estimated to be about 13.5–17.5 m (44–57 ft), almost similar to that of the modern sperm whale (Physeter macrocephalus), making it one of the largest predators known to have existed. The teeth of Livyatan measured 36.2 cm (1.19 ft), and are the largest biting teeth of any known animal, excluding tusks. It is distinguished from the other raptorial sperm whales by the basin on the skull spanning the length of the snout. The spermaceti organ contained in that basin is thought to have been used in echolocation and communication, or for ramming prey and other sperm whales. The whale may have interacted with the large extinct shark megalodon (Otodus megalodon), competing with it for a similar food source. Its extinction was probably caused by a cooling event at the end of the Miocene period causing a reduction in food populations. The geological formation where the whale has been found has also preserved a large assemblage of marine life, such as sharks and marine mammals. (Full article...) -
Rice's whale (Balaenoptera ricei), also known as the Gulf of Mexico whale, is a species of baleen whale endemic to the northern Gulf of Mexico. Initially identified as a subpopulation of the Bryde's whale, genetic and skeletal studies found it to be a distinct species by 2021. In outward appearance, it is virtually identical to the Bryde's whale. Its body is streamlined and sleek, with a uniformly dark charcoal gray dorsal and pale to pinkish underside. A diagnostic feature often used by field scientists to distinguish Rice's whales from whales other than the Bryde's whale is the three prominent ridges that line the top of its head. The species can be distinguished from the Bryde's whale by the shape of the nasal bones, which have wider gaps due to a unique wrapping by the frontal bones, its unique vocal repertoire, and genetic differences.
It is a medium-sized baleen whale that grows up to 12.65 meters (41.5 ft) in length and weighs up to 13.87–27.2 metric tons (13.65–26.77 long tons; 15.29–29.98 short tons). The Rice's whale inhabits a restricted stretch along the continental slope in the northeastern part of the Gulf of Mexico between depths of 150–410 meters (490–1,350 ft) off the coast of western Florida, although some whales have been sighted in the northwestern portions and the species may have inhabited a wider distribution throughout the Gulf in historical times. It does not migrate but remains within this area year-round. Little is known about the feeding behavior of Rice's whales, but data from a tagged individual revealed a diel vertical diving pattern, in which the whale spends most of the day feeding at or near the seafloor at depths of up to 271 meters (889 ft) and night at the surface. The whale's diet remains unknown, but lanternfish and hatchetfish are suspected prey. (Full article...) -
Pilot whales are cetaceans belonging to the genus Globicephala. The two extant species are the long-finned pilot whale (G. melas) and the short-finned pilot whale (G. macrorhynchus). The two are not readily distinguishable at sea, and analysis of the skulls is the best way to distinguish between the species. Between the two species, they range nearly worldwide, with long-finned pilot whales living in colder waters and short-finned pilot whales living in tropical and subtropical waters. Pilot whales are among the largest of the oceanic dolphins, exceeded in size only by the orca. They and other large members of the dolphin family are also known as blackfish.
Pilot whales feed primarily on squid, but will also hunt large demersal fish such as cod and turbot. They are highly social and may remain with their birth pod throughout their lifetime. Short-finned pilot whales are one of the few non-primate mammal species in which females go through menopause, and postreproductive females continue to contribute to their pod. Pilot whales are notorious for stranding themselves on beaches, but the reason behind this is not fully understood. Marine biologists have shed some light on the matter, suggesting that it is due to the mammals inner ear (their principal navigational sonar) being damaged from noise pollution in the ocean, such as from cargo ships or military exercises. The conservation status of short-finned and long-finned pilot whales has been determined to be least concern. (Full article...) -
Granny (born c. 1936-1951 approx, died c. 2016), also known as J2, was a female orca of the J pod of southern resident orcas notable for her long life. Early estimates placed her birth in 1911, putting her at 105 years old at the time of her death. However, this estimate was later theorized to have been based on mistaken information and more recent studies put her at 65–80 years old. If she was 105, she would have been the oldest known orca at the time of her death. Granny lived in the northeast Pacific Ocean and coastal bays of Washington state and British Columbia. She was last seen on October 12, 2016, and was considered deceased by The Center for Whale Research in January 2017. (Full article...) -
The dwarf sperm whale (Kogia sima) is a sperm whale that inhabits temperate and tropical oceans worldwide, in particular continental shelves and slopes. It was first described by biologist Richard Owen in 1866, based on illustrations by naturalist Sir Walter Elliot. The species was considered to be synonymous with the pygmy sperm whale (Kogia breviceps) from 1878 until 1998. The dwarf sperm whale is a small whale, 2 to 2.7 m (6 ft 7 in to 8 ft 10 in) and 136 to 272 kg (300 to 600 lb), that has a grey coloration, square head, small jaw, and robust body. Its appearance is very similar to the pygmy sperm whale, distinguished mainly by the position of the dorsal fin on the body–nearer the middle in the dwarf sperm whale and nearer the tail in the other.
The dwarf sperm whale is a suction feeder that mainly eats squid, and does this in small pods of typically one to four members. It is preyed upon by the killer whale (Orcinus orca) and large sharks such as the great white shark (Carcharodon carcharius). When startled, the whale can eject a cloud of red-brown fluid. Most of what is known of the whale comes from beached individuals, as sightings in the ocean are rare. Many of these stranded whales died from parasitic infestations or heart failure. (Full article...) -
Baleen whales (/bəˈliːn/), also known as whalebone whales, are marine mammals of the parvorder Mysticeti in the infraorder Cetacea (whales, dolphins and porpoises), which use keratinaceous baleen plates (or "whalebone") in their mouths to sieve planktonic creatures from the water. Mysticeti comprises the families Balaenidae (right and bowhead whales), Balaenopteridae (rorquals), Eschrichtiidae (the gray whale) and Cetotheriidae (the pygmy right whale). There are currently 16 species of baleen whales. While cetaceans were historically thought to have descended from mesonychians, molecular evidence instead supports them as a clade of even-toed ungulates (Artiodactyla). Baleen whales split from toothed whales (Odontoceti) around 34 million years ago.
Baleen whales range in size from the 6 m (20 ft) and 3,000 kg (6,600 lb) pygmy right whale to the 31 m (102 ft) and 190 t (210 short tons) blue whale, the largest known animal to have ever existed. They are sexually dimorphic. Baleen whales can have streamlined or large bodies, depending on the feeding behavior, and two limbs that are modified into flippers. The fin whale is the fastest baleen whale, recorded swimming at 10 m/s (36 km/h; 22 mph). Baleen whales use their baleen plates to filter out food from the water by either lunge-feeding or skim-feeding. Baleen whales have fused neck vertebrae, and are unable to turn their heads at all. Baleen whales have two blowholes. Some species are well adapted for diving to great depths. They have a layer of fat, or blubber, under the skin to keep warm in the cold water. (Full article...) -
Whales are a widely distributed and diverse group of fully aquatic placental marine mammals. As an informal and colloquial grouping, they correspond to large members of the infraorder Cetacea, i.e. all cetaceans apart from dolphins and porpoises. Dolphins and porpoises may be considered whales from a formal, cladistic perspective. Whales, dolphins and porpoises belong to the order Cetartiodactyla, which consists of even-toed ungulates. Their closest non-cetacean living relatives are the hippopotamuses, from which they and other cetaceans diverged about 54 million years ago. The two parvorders of whales, baleen whales (Mysticeti) and toothed whales (Odontoceti), are thought to have had their last common ancestor around 34 million years ago. Mysticetes include four extant (living) families: Balaenopteridae (the rorquals), Balaenidae (right whales), Cetotheriidae (the pygmy right whale), and Eschrichtiidae (the grey whale). Odontocetes include the Monodontidae (belugas and narwhals), Physeteridae (the sperm whale), Kogiidae (the dwarf and pygmy sperm whale), and Ziphiidae (the beaked whales), as well as the six families of dolphins and porpoises which are not considered whales in the informal sense.
Whales are fully aquatic, open-ocean animals: they can feed, mate, give birth, suckle and raise their young at sea. Whales range in size from the 2.6 metres (8.5 ft) and 135 kilograms (298 lb) dwarf sperm whale to the 29.9 metres (98 ft) and 190 tonnes (210 short tons) blue whale, which is the largest known animal that has ever lived. The sperm whale is the largest toothed predator on Earth. Several whale species exhibit sexual dimorphism, in that the females are larger than males. (Full article...) -
Zygophyseter varolai is an extinct sperm whale that lived during the Tortonian age of the Late Miocene 11.2 to 7.6 million years ago. It is known from a single specimen from the Pietra Leccese Formation in Italy. It was a member of a stem group of fossil macroraptorial sperm whales (often shortened to "raptorial") also including Brygmophyseter, Acrophyseter, and Livyatan. It probably grew to be around 6.5 to 7 meters (21 to 23 ft) in length and shared some characteristics with other raptorials, such as large teeth with tooth enamel that were functional in both the upper and lower jaws which the modern sperm whale (Physeter macrocephalus) lacks. It also had a beak, the ability to echolocate prey, and could have probably swum faster than the modern-day sperm whale which can reach 4 kilometers per hour (2.5 mph). These were probably used in the capture of large prey, such as large fish, seals, and whales. In fact, its common name, the killer sperm whale, refers to its feeding habits that would have had a resemblance to the modern-day killer whale (Orcinus orca). (Full article...) -
Ambulocetus (Latin ambulare "to walk" + cetus "whale") is a genus of early amphibious cetacean from the Kuldana Formation in Pakistan, roughly 48 or 47 million years ago during the Early Eocene (Lutetian). It contains one species, Ambulocetus natans (Latin natans "swimming"), known solely from a near-complete skeleton. Ambulocetus is among the best-studied of Eocene cetaceans, and serves as an instrumental find in the study of cetacean evolution and their transition from land to sea, as it was the first cetacean discovered to preserve a suite of adaptations consistent with an amphibious lifestyle. Ambulocetus is classified in the group Archaeoceti—the ancient forerunners of modern cetaceans whose members span the transition from land to sea—and in the family Ambulocetidae, which includes Himalayacetus and Gandakasia (also from the Eocene of the Indian subcontinent).
Ambulocetus had a narrow, streamlined body, and a long, broad snout, with eyes positioned at the very top of its head. Because of these features, it is hypothesised to have behaved much like a crocodile, waiting near the water's surface to ambush large mammals, using its powerful jaws to clamp onto and drown or thrash prey. Additionally, its ears possessed similar traits to modern cetaceans, which are specialised for hearing and detecting certain frequencies underwater, although it is unclear if Ambulocetus also used these specialised ears for hearing underwater. They may have instead been utilised for bone conduction on land, or perhaps served no function for early cetaceans. (Full article...) -
Orcinus meyeri is a fossil species of Orcinus (killer whales) found in the Early Miocene deposits of southern Germany, known from two jaw fragments and 18 isolated teeth. It was originally described as Delphinus acutidens in 1859, but reclassified in 1873. Its validity is disputed, and it may be a synonymous with the ancient sperm whale Physeterula dubusi. It was found in the Alpine town of Stockach in the Molasse basin, which was a coastal area with strong tidal currents. (Full article...) -
The sperm whale or cachalot (Physeter macrocephalus) is the largest of the toothed whales and the largest toothed predator. It is the only living member of the genus Physeter and one of three extant species in the sperm whale family, along with the pygmy sperm whale and dwarf sperm whale of the genus Kogia.
The sperm whale is a pelagic mammal with a worldwide range, and will migrate seasonally for feeding and breeding. Females and young males live together in groups, while mature males (bulls) live solitary lives outside of the mating season. The females cooperate to protect and nurse their young. Females give birth every four to twenty years, and care for the calves for more than a decade. A mature, healthy sperm whale has no natural predators, although calves and weakened adults are sometimes killed by pods of killer whales (orcas). (Full article...) -
Boudewijn Seapark dolphinarium in Bruges, Belgium
A dolphinarium is an aquarium for dolphins. The dolphins are usually kept in a pool, though occasionally they may be kept in pens in the open sea, either for research or public performances. Some dolphinariums (sometimes called dolphinaria in plural) consist of one pool where dolphins perform for the public, others are part of larger parks, such as marine mammal parks, zoos or theme parks, with other animals and attractions as well.
While cetaceans have been held in captivity since the 1860s, the first commercial dolphinarium was opened only in 1938. Their popularity increased rapidly until the 1960s. Since the 1970s, increasing concern for animal welfare led to stricter regulation, which in several countries ultimately resulted in the closure of some dolphinariums. Despite this trend, dolphinariums are still widespread in Europe, Japan and North America. The most common species of dolphin kept in dolphinariums is the bottlenose dolphin, as it is relatively easy to train and has a long lifespan in captivity. While trade in dolphins is internationally regulated, other aspects of keeping dolphins in captivity, such as the minimum size and characteristics of pools, vary among countries. Though animal welfare is perceived to have improved significantly over the last few decades, many animal rights groups still consider keeping dolphins captive to be a form of animal abuse. (Full article...)
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Photo credit: U.S. Navy photo by Photographer's Mate 1st Class Brien Aho.
A U.S. Navy Marine Mammal Program bottlenose dolphin named K-Dog wearing a locating pinger, performed mine clearance work in the Persian Gulf during the Iraq War.
The United States and Russian militaries have trained and employed dolphins for several reasons. Such military dolphins can be trained to rescue lost divers or to locate underwater mines. Military dolphins were used during the First and Second Gulf War.
More did you know...
- ...common dolphins, which are often seen off South Africa’s east coast, can occur in schools of several thousand. The biggest school on record was estimated to consist of about 15,000 dolphins!
- ...because whales and dolphins are streamlined to swim in water, they do not have external organs. This makes it almost impossible to tell the sex of a whale or dolphin when watching them on the sea surface.
- ...there are probably types of cetaceans that are as yet unknown. For example, the Longman's beaked whale is only known from skulls washed ashore in Somalia and Australia. It has never been seen alive!
- ...in baleen whales females are bigger than males, while in the toothed whales the males are bigger than females.
- ...the open mouth of an adult southern right whale can be two metres wide!
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General images -
The following are images from various cetacean-related articles on Wikipedia.
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Archaeomysticetes, like Janjucetus, had teeth. -
Bryde's whale (rorqual) (from Baleen whale) -
Restoration of Janjucetus hunderi (from Baleen whale) -
Antarctic minke whale illustration -
A phylogenetic tree showing the relationships among cetacean families. (from Evolution of cetaceans) -
Megalodon hunted Miocene baleen whales (from Baleen whale) -
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Dolphins (aquatic mammals) and ichthyosaurs (extinct marine reptiles) share a number of unique adaptations for fully aquatic lifestyle and are frequently used as extreme examples of convergent evolution (from Evolution of cetaceans) -
Features of a sperm whale skeleton (from Toothed whale) -
Common minke whale illustration -
The heart of a blue whale with a person standing next to it -
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Fin whale illustration -
Bowhead whale illustration -
The remains of a North Atlantic right whale after it collided with a ship propeller -
The skeleton of a bowhead whale with the hind limb and pelvic bone structure circled in red. This bone structure stays internal during the entire life of the species. (from Evolution of cetaceans) -
Diagram illustrating sound generation, propagation and reception in a toothed whale. Outgoing sounds are red and incoming ones are green (from Toothed whale) -
Atlantic white-sided dolphin caught in a drive hunt in Hvalba on the Faroe Islands being taken away with a forklift (from Toothed whale) -
Fossil of Squalodon (from Toothed whale) -
A protest against Japan's scientific whaling -
Baleen whales vary considerably in size and shape, depending on their feeding behavior. (from Baleen whale) -
Spectrogram of dolphin vocalizations. Whistles, whines, and clicks are visible as upside down V's, horizontal striations, and vertical lines, respectively. (from Toothed whale) -
Eden's whale illustration -
Accessory baleen plates taper off into small hairs (from Baleen whale) -
A humpback whale skeleton. Notice how the jaw is split into two. -
An orca by the name of Ulises performing at SeaWorld, 2009 (from Toothed whale) -
Humpback whales lunge-feeding in the course of bubble net fishing -
Acrophyseter skull (from Evolution of cetaceans) -
The nose of the whale is filled with a waxy substance that was widely used in candles, oil lamps, and lubricants (from Toothed whale) -
Biosonar by cetaceans (from Toothed whale) -
Pygmy right whale illustration -
Japanese scientific whaling on a pair of Antarctic minke whales (from Baleen whale) -
Orange whale lice on a right whale -
Species of the infraorder Cetacea (from Evolution of cetaceans) -
Anatomy of the bottlenose dolphin (from Toothed whale) -
Pakicetus attocki skeleton (from Evolution of cetaceans) -
Blue whale illustration -
World population graph of blue whales -
Orca hunting a Weddell seal (from Toothed whale) -
South Atlantic right whale illustration -
Female right whale with calf -
Humpback whale illustration -
Possible relationships between cetaceans and other ungulate groups. (from Evolution of cetaceans) -
Archaeocetes (like this Basilosaurus) had a heterodont dentition (from Evolution of cetaceans) -
Sei whale illustration -
Their eyes are relatively small for their size. -
North Atlantic right whale illustration -
Gray whale illustration
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Did you know (auto-generated)
- ... that the South Asian river dolphin is nearly blind and relies on echolocation for navigation?
- ... that one of the first researchers to propose dolphin-assisted therapy for humans later renounced it?
- ... that Celia Kaye won the Golden Globe Award for Most Promising Newcomer in 1965 for her starring role in Island of the Blue Dolphins?
- ... that one can swim with humpback whales in the Niue Nukutuluea Multiple-Use Marine Park?
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